COLEOPTERA - SCARABAEIDAE - SCARABAEINAE

As soon as we know how to divide the huge list below into the five tribes, this will be easier to use.

Canthonini
Coprini
Onitini
Onthaphagini
Scarabaeini

Agamopus castaneus Balthasar
Agamopus lampros Bates 1887

Allogymnopleurus aeneas (Harold) 1875
Allogymnopleurus olivieri (Laporte) 1840
Allogymnopleurus thallassinus (Klug) 1855
Allogymnopleurus umbrinus (Gerstaecker) 1871

Amphistomus speculifer Matthews

Anachalcos convexus Boheman 1857

Anisocanthon villosum (Harold) 1868

Anomiopsoides heteroclyta (Blanchard) 1843

Anonychonitis freyi

Anoplodrepanus pecki (Howden)
Anoplodrepanus reconditus (Matthews) 

Aphengium sordidum Harold, 1868

Aptenocanthon hopsini Carter

Ateuchus aeneomicans (Harold) 1868
Ateuchus ampliatus (Bates) 1888
Ateuchus candezei (Harold)
Ateuchus halffteri Kohlmann
Ateuchus histeroides (Weber) 1801
Ateuchus lecontei (Harold) 1867
Ateuchus opacipennis (Fairmaire) 1884
Ateuchus rodriguezi (DeBorre)
Ateuchus texanus Robinson

Bdelyropsis bowditchi (Paulian) 1938
Bdelyrus seminudum (Bates) 1887

Bolbites onitoides Harold 1868

Bubas bison Linnaeus 1767
Bubas bubalus Olivier 1811

Caccobius denticollis Harold 1897
Caccobius ferrugineus Fahraeus 1857
Caccobius gonoderus Fairmaire 1888
Caccobius meridionalis Boucomont 1914
Caccobius schreberi (Linnaeus) 1767
Caccobius ultor Sharp 1875
Caccobius vulcanus Fabricius 1867

Canthidium ardens Bates 1887
Canthidium aurifex Bates 1887
Canthidium breve (Germar) 1824
Canthidium centrale Boucomont 1928
Canthidium elegantulum Balthasar
Canthidium epistomale Boucomont 1928
Canthidium haroldi Preudhomme 1886
Canthidium hespenheidei Howden & Young 1981
Canthidium leucopterum Howden & Young 1981
Canthidium pallidoalatum Howden & Young 1981
Canthidium perceptibile Howden & Young 1981
Canthidium puncticolle Harold 1867
Canthidium tuberifrons Howden & Young 1981
Canthidium vespertinum Howden & Young 1981
Canthochilum hispidum serropunctata

Canthon acutoides Schmidt 1922
Canthon acutus Harold 1868
Canthon aequinoctialis Harold 1868
Canthon aereum (Schmidt) 1922
Canthon affinis (Fabricius) 1801
Canthon angularis Harold 1868
Canthon angustatus Harold 1869
Canthon baltheatus Boheman 1858
Canthon bispinatus Robinson
Canthon chalcites (Haldeman) 1843
Canthon chevrolati Harold 1868
Canthon conformis Harold 1868
Canthon curvodilatatus Schmidt
Canthon cyanellus LeConte 1859
Canthon cyanocephalus Harold 1868
Canthon delicatulus Balthasar 1939
Canthon depressipennis LeConte 1859
Canthon deyrollei Harold 1868
Canthon ebenus (Say) 1823
Canthon edentulus Harold 1868
Canthon euryscelis Bates 1887
Canthon femoralis Chevrolat 1834
Canthon femoralis balsensis
Canthon fortemarginatus Balthasar 1939
Canthon fulgidus Redtenbacher 1867
Canthon fuscipes Erichson 1847
Canthon hidalgoensis Bates 1887
Canthon humectus (Say) 1832
Canthon imitator Brown 1946
Canthon indigaceus LeConte 1866
Canthon indigaceus chevrolati Harold 1868
Canthon juvencus Harold 1868
Canthon lamprimus Bates 1887
Canthon latipes Blanchard 1843
Canthon lecontei Harold 1868
Canthon literatus Germar 1813
Canthon lividus Blanchard 1843
Canthon luteicollis Erichson 1847
Canthon melanus Robinson 1948
Canthon monilifer Blanchard 1843
Canthon morsei Howden 1966
Canthon mutabilis Lucas 1857
Canthon nigricornis Say 1823
Canthon nigripennis Lansberg 1874
Canthon opacum Lucas 1857
Canthon pallidus Schmidt 1922
Canthon perplexum LeConte 1847
Canthon pilularius (Linnaeus) 1758
Canthon politus Harold 1868
Canthon praticola LeConte 1859 
Canthon probus Germar, 1824
Canthon puncticollis LeConte 1866
Canthon quadriguttatus Olivier 1789
Canthon quinquemaculatus Laporte 1840
Canthon quinqueplicatus? Laporte
Canthon rubrescens Blanchard 1843
Canthon scrutator Balthasar 1939
Canthon septemmaculatus (Latreille) 1807
Canthon simplex LeConte 1857
Canthon simplex militaris Horn 1878
Canthon smaragdulus (Fabricius) 1781
Canthon sordidus Harold 1868
Canthon subhyalinus Harold 1867
Canthon tetraodor Blanchard 1843
Canthon triangularis (Drury) 1770
Canthon vigilans LeConte 1858
Canthon violaceus Olivier 1789
Canthon virens (Mannerheim) 1829
Canthon viridis meridionalis Martinez & Halfftler 1964

Catharsius capucinus Fabricius 1781
Catharsius dux Harold 1878
Catharsius furcillatus Guerin 1847
Catharsius guineensis Ferreira
Catharsius molossus Linnaeus 1758
Catharsius mossambicanus Ferreira 1967
Catharsius pithecius (Fabricius) 1775
Catharsius platycerus Klug 1855
Catharsius sagax Quens. 1806
Catharsius sesostris Waterhouse 1888
Catharsius ulysses Boheman 1857

Cephalodesmius armiger Westwood 1841

Chalcocopris hespera (Olivier, 1789)

Chironitis furcifer Rossi 1792
Chironitis hungaricus (Herbst) 1789
Chironitis irorratus (Rossi) 1790
Chironitis irroratus (Rossi) 1790
Chironitis pamphilus Menetries 1849
Chironitis scabrosus Fabricius 1776

Circellium bacchus (Fabricius) 1781

Copris acutidens Motschulsky 1860
Copris amyntor Klug 1855
Copris arizonensis Schaeffer 1906
Copris boucardi Harold 1869
Copris costaricensis Gahan 1894
Copris delicatus Arrow
Copris elphenor Klug 1855
Copris fallaciosus Gillet 1907
Copris fricator Fabricius 1787
Copris hintoni Saylor 1933
Copris hispanus Linnaeus 1764
Copris howdeni Matthews
Copris imitans Felsche 1910
Copris incertus incertus Say 1835
Copris inemarginatus Blatchley 1918
Copris interioris Kolbe
Copris klugi Harold 1869
Copris laeviceps Harold 1869
Copris laius Harold 1868
Copris lecontei Matthews 1962
Copris lugubris Boheman 1858
Copris lunaris (Linnaeus) 1758
Copris macclevei Warner
Copris mesacanthus Harold 1878
Copris minutus Drury 1773
Copris ochus (Motschulsky) 1860
Copris pueli Boissy 1905
Copris rebouchei Harold 1869
Copris remotus LeConte 1866
Copris signatus Walker 1858
Copris sinon Olivier 1789
Copris subsidens Peringuey 1900
Copris umbilicatus Abeille de Perrin
Copris victorini Boheman 1857

Coprophanaeus chiriquensis Olson
Coprophanaeus dardanus MacLeay 1819
Coprophanaeus jasius (Olivier) 1789 
Coprophaneus morenoi Arnaud
Coprophanaeus parvulus Olsoufieff 1924
Coprophanaeus pecki Howden & Young 1981
Coprophanaeus pluto (Harold) 1863
Coprophanaeus saphirinus Sturm 1826
Coprophanaeus spitzi Pessoa 1934
Coprophanaeus telamon (Erichson) 1847

Coptodactyla brooksi Matt.
Coptodactyla glabricollis Hope 1841
Coptodactyla onitoides Gyllenhal
Coptodactyla torresita

Coptorhina auspicata Peringuey 1901
Coptorhina klugi Hope 1830

Cryptocanthon nebulinus Howden 1973
Cryptocanthon parvus Howden

Cyptochirus ambiguus (Kirby) 1828

Deltochilum amazonicum Bates 1887
Deltochilum brasiliense Laporte 1840
Deltochilum carinatum Westwood 1837
Deltochilum femorale Bates 1870
Deltochilum furcatum Laporte 1840
Deltochilum gibbosum (Fabricius) 1775
Deltochilum guyanense Paulian 1933
Deltochilum icarus Olivier 1789
Deltochilum lobipes Bates 1887
Deltochilum mexicanum Burmeister 1898
Deltochilum morbillosum Burmeister 1848
Deltochilum orbiculare Lansberg 1874
Deltochilum orbignyi Blanchard 1843
Deltochilum parile Bates 1887
Deltochilum pseudoparile Paulian 1938
Deltochilum scabrisculum Bates 1887
Deltochilum septemstriatum Paulian 1938
Deltochilum valgum Burmeister 1873

Dendropaemon denticollis Felsche 1909
Dendropaemon hirticollis D'Olsoufieff 1924

Diabroctis mimas (Linnaeus) 1758

Diastellopalpus dudleyi Cambefort
Diastellopalpus fuelleborni (Kolbe) 1900
Diastellopalpus infernalis Lansberg 1887
Diastellopalpus johnstoni (Waterhouse) 1885
Diastellopalpus neavei d'Orbigny
Diastellopalpus quinquedens Bates 1888
Diastellopalpus sulciger Kolbe 1893
Diastellopalpus thomsoni (Bates) 1888
Diastellopalpus tridens (Fabricius) 1781

Dichotomius achamas (Harold) 1867
Dichotomius agenor (Harold) 1869
Dichotomius amplicollis (Harold) 1869
Dichotomius anaglypticus (Mannerheim) 1829
Dichotomius ascanius (Harold) 1869
Dichotomius assifer (Eschscholtz) 1822
Dichotomius batesi (Harold) 1869
Dichotomius boreus (Olivier) 1789
Dichotomius bosquei (Periera)
Dichotomius carbonarius (Mannerheim) 1829
Dichotomius carolinus (Linnaeus) 1767
Dichotomius centralis (Harold) 1869
Dichotomius colonicus colonicus (Say) 1835
Dichotomius conicollis (Blanchard) 1843
Dichotomius gamboaensis Howden & Young 1981
Dichotomius horrida (Felsche) 1911
Dichotomius luctuosus (Harold) 1869
Dichotomius mormon Ljungh 1799
Dichotomius nisus (Olivier) 1789
Dichotomius nobilis (Waterhouse) 1891
Dichotomius protectus (Harold) 1867
Dichotomius quadraticeps (Felsche) 1901
Dichotomius satanus (Harold) 1867
Dichotomius semiaeneus (Germar) 1824
Dichotomius telamon (Harold) 1869
Dichotomius torulosus (Eschscholtz) 1822
Dichotomius yucatanus (Bates) 1887

Diorgopyx asciculiferum Matthews Diorgopyx incrassifer

Drepanocerus abyssinicus Roth 1851
Drepanocerus caelatus (Gerstaecker) 1871
Drepanocerus fastiditus Peringuey 1901
Drepanocerus freyi Janssens 1953
Drepanocerus impressicollis Boheman 1857
Drepanocerus kirbyi Kirby 1828
Drepanocerus laticollis Fahraeus 1857
Drepanocerus marshalli Boucomont 1921
Drepanocerus patrizii Boucomont 1923
Drepanocerus setosus Wiedemann 1823
Drepanocerus sulcicollis Laporte 1840
Drepanoplatys gilleti Boucomont 1921
Drepanopodus proximus (Peringuey) 1901

Epirinus flagellatus (Fabricius) 1775
Epirinus silvestris (Cambefort)
Epirinus validus Peringuey 1901

Euoniticellus africanus (Harold) 1873
Euoniticellus fulvus (Goeze) 1777
Euoniticellus inaequalis (Reiche) 1850
Euoniticellus intermedius (Reiche) 1849
Euoniticellus kawanus (Janssens) 1939
Euoniticellus pallens (Olivier) 1789
Euoniticellus pallipes (Fabricius) 1781
Euoniticellus triangulatus (Harold) 1873
Euoniticellus zumpti Janssens 1953

Eurysternus angustulus Harold 1869
Eurysternus caribaeus Herbst 1789
Eurysternus confusus Jessop
Eurysternus deplanatus Guerin 1824
Eurysternus foedus Kirsch 1871
Eurysternus magnus Laporte 1840
Eurysternus marmoreus Laporte 1840
Eurysternus mexicanus Harold 1869
Eurysternus parallelus Laporte 1840
Eurysternus plebejus Harold 1880
Eurysternus velutinus Bates 1887

Garreta azureus (Fabricius) 1801
Garreta malleolus (Kolbe) 1895
Garreta nitens (Olivier) 1789
Garreta unicolor (Fahraeus) 1857

Glaphyrocanthon variabilis Martinez Glaphyrocanthon viridis (Beauvois) 1805

Goniocanthon smaragdinum (Fabricius) 1781

Gromphas aeruginosa (Perty) 1830 Gromphus lacordairei Brulle 1834

Gymnopleurus asperrimus Felsche 1909
Gymnopleurus cyaneus (Fabricius) 1798
Gymnopleurus flagellatus (Fabricius) 1787
Gymnopleurus fulgidus (Olivier) 1789
Gymnopleurus gemmatus Harold 1871
Gymnopleurus geoffroyi (Fuessly) 1775
Gymnopleurus humanus MacLeay 1821
Gymnopleurus ignitis Klug 1855
Gymnopleurus koenigi (Fabricius) 1775
Gymnopleurus melanarius Harold 1867
Gymnopleurus miliaris (Fabricius) 1775
Gymnopleurus modestus Peringuey 1888
Gymnopleurus mopsus (Pallas) 1781
Gymnopleurus nyankpalaensis Endrodi
Gymnopleurus sericeifrons Fairmaire 1887
Gymnopleurus sinuatus (Olivier) 1789
Gymnopleurus sturmi MacLeay 1821
Gymnopleurus thewalli Waterhouse 1891
Gymnopleurus tristis Laporte 1840
Gymnopleurus virens Erichson 1843

Gyronotus marginatus Peringuey 1888

Hansreia affinis (Fabricius) 1801

Helictopleurus giganteus (Harold) 1869
Helictopleurus marsyas (Olivier) 1789
Helictopleurus quadripunctatus (Olivier) 1789

Heliocopris andersoni Bates 1868
Heliocopris antenor (Olivier) 1789
Heliocopris bucephalus (Fabricius) 1775
Heliocopris dominus Bates 1868
Heliocopris hamadryas (Fabricius) 1775
Heliocopris hamifer Harold 1878
Heliocopris isidis Latreille 1819
Heliocopris japetus Klug 1855
Heliocopris neptunus Boheman 1857
Heliocopris pirmal (Fabricius) 1798
Heliocopris staudingeri Kolbe 1893

Heteronitis castelnaui (Harold) 1862

Hyalonthophagus alcyon (Klug)

Kheper aegyptiorum (Latreille) 1827
Kheper cupreus (Laporte) 1840
Kheper erichsoni (Harold) 1867
Kheper lamarcki (MacLeay) 1821
Kheper nigroaenus (Boheman) 1857
Kheper platynotus (Bates) 1888
Kheper prodigiosus (Erichson) 1843
Kheper subaeneus (Harold) 1869

Lepanus australis Matthews
Lepanus illawarrensis Matthews 1974

Liatongus affinis (Arrow) 1908
Liatongus arrowi (Boucomont) 1921
Liatongus californicus (Horn) 1882
Liatongus femoralis (Illiger) 1800
Liatongus gagatinus (Hope) 1831
Liatongus indicus Arrow 1908
Liatongus margaceras (Hope) 1831
Liatongus militaris (Castelnau) 1840
Liatongus monstrosus (Bates) 1887
Liatongus phanaeoides (Westwood) 1839
Liatongus rhadamistus (Fabricius) 1775
Liatongus rhinocerulus (Bates) 1889
Liatongus taurus (Boucumont) 1920
Liatongus venator (Fabricius) 1801

Madateuchus viettei Paulian 1953

Malagionella astynax yucateca (Harold) 1863

Malagionella bicolor Guerin 1964

Megaphanaeus bonariensis (Gory) 1844
Megaphanaeus ensifer (Germar) 1821
Megaphanaeus lancifer (Linnaeus) 1758

Megatharsis buckleyi Waterhouse 1891

Megathopa aeneicollis Waterhouse 1890
Megathopa argentina Gillet 1911
Megathopa villosa Eschscholtz 1822
Megathopa violacea Blanchard 1843

Megathoposoma candezei Harold 1873

Melanocanthon granulifer (Schmidt)

Mentophilus hollandiae Laporte 1792

Milichus apicalis (Fahraeus) 1857

Monoplistes leai Paulian

Neateuchus proboscideus (Guerin) 1844

Neopachysoma denticolle (Periguey) 1888
Neopachysoma rotundigena (Felsche) 1907

Neosisyphus spinipes (Thunberg) 1818

Odontoloma louwi Howden & Scholtz 1987

Oniticellus cinctus (Fabricius) 1775
Oniticellus egregius Klug 1885
Oniticellus pictus (Hausman) 1807
Oniticellus planatus Laporte 1840

Onitis aerarius Harold 1878
Onitis alexis Klug 1835
Onitis aygulus (Fabricius) 1781
Onitis belial Fabricius 1789
Onitis consanguineus Felsche
Onitis crenatus Reiche 1850
Onitis curvipes Lansberge 1875
Onitis deceptor Peringuey 1901
Onitis falcatus (Wulfen) 1789
Onitis fulgidus Klug 1855
Onitis guineensis Gillet 1918
Onitis humerosus (Pallas) 1771
Onitis inversidens Lansberge 1875
Onitis ion (Olivier) 1789
Onitis niger Lansberge 1875
Onitis numida Laporte 1840
Onitis obenbergeri Balthasar 1942
Onitis phartopus Lansberge 1875
Onitis philemon Fabricius 1801
Onitis picticollis Boheman 1857
Onitis reichei Lansberge 1875
Onitis subopacus Arrow 1931
Onitis uncinatus Klug 1855
Onitis violaceus Lansberge 1875
Onitis virens Lansberge 1875
Onitis viridulus Boheman 1857
Onitis westermanni Lansberge 1886

Ontherus alexis (Blanchard) 1845
Ontherus appendiculatus (Mannerheim) 1829
Ontherus brevipennis Harold 1867
Ontherus didymus Erichson 1847
Ontherus edentulus Genier 1992
Ontherus mexicanus Harold 1869
Ontherus pseudodidymus Genier 1992
Ontherus pubens Genier 1993
Ontherus sulcator (Fabricius) 1775
Ontherus ulcopygus Genier 1992

Onthocharis panamensis Paulian 1939

Onthophagus acuminatus Harold 1880
Onthophagus acutus d'Orbigny 1908
Onthophagus aeruginosus Roth 1851
Onthophagus albipodex d'Orbigny 1902
Onthophagus alluvius Howden & Cartwright
Onthophagus amphinasus Arrow 1931
Onthophagus amphioxus Arrow
Onthophagus amyntus (Olivier) 1789
Onthophagus andalusicus Waltl 1835
Onthophagus arilla Matthews 1972
Onthophagus arnetti Howden & Cartwright 1963
Onthophagus atrox Harold 1867
Onthophagus australis Guerin 1830
Onthophagus batesi Howden & Cartwright 1963
Onthophagus belorhinus Bates 1887
Onthophagus bicavifrons d'Orbigny 1902
Onthophagus bidens (Olivier) 1789
Onthophagus bifasciatus (Fabricius) 1781
Onthophagus binodis Thunberg 1818
Onthophagus bituberculatus (Olivier) 1789
Onthophagus bonasus (Fabricius) 1775
Onthophagus bovinus Peringuey 1892
Onthophagus bronzaeus Arrow 1907
Onthophagus browni Howden & Cartwright 1963
Onthophagus cameloides d'Orbigny 1900
Onthophagus capella Kirby 1818
Onthophagus carbonarius Klug 1855
Onthophagus castetsi Lansberge 1887
Onthophagus cervus (Fabricius) 1798
Onthophagus championi Bates 1887
Onthophagus cinctipennis Quedenfeldt 1884
Onthophagus cochisus Brown 1927
Onthophagus coenobita (Herbst) 1783
Onthophagus concinnus Laporte
Onthophagus consentaneus Harold 1867
Onthophagus coproides Horn 1881
Onthophagus coscineus Bates 1887
Onthophagus cribellum d'Orbigny 1902
Onthophagus cribricollis Horn 1881
Onthophagus cribripennis d'Orbigny 1902
Onthophagus crinitus Harold 1869
Onthophagus crinitus panamensis Bates 1887
Onthophagus curvicornis curvicornis Latreille 1811
Onthophagus curvicornis incensus Say 1837
Onthophagus cyanellus Bates 1887
Onthophagus dama (Fabricius) 1798
Onthophagus dandalus Matthews 1972
Onthophagus depressus Harold 1871
Onthophagus deterrans Peringuey 1901
Onthophagus diabolicus Harold 1877
Onthophagus dicranius Bates 1887
Onthophagus duporti Boucomont 1914
Onthophagus ebenicolor d'Orbigny 1902
Onthophagus ebenus Peringuey 1888
Onthophagus emeritus Peringuey 1901
Onthophagus ensifer Boucomont 1914
Onthophagus ephippioderus Arrow 1907
Onthophagus fallax d'Orbigny 1913
Onthophagus fasciatus Boucomont 1914
Onthophagus ferox Harold 1867
Onthophagus fimetarius Roth 1851
Onthophagus fissiceps MacLeay 1888
Onthophagus flavimargo d'Orbigny 1902
Onthophagus flavolimbatus Klug 1855
Onthophagus foveolatus Harold 1877
Onthophagus fracticornis (Preyssler) 1790
Onthophagus furcatus Fabricius 1781
Onthophagus furcillifer Bates 1891
Onthophagus fuscus Boucomont 1914
Onthophagus gazella Fabricius 1787
Onthophagus granulatus Boheman 1858
Onthophagus granulum d'Orbigny 1913
Onthophagus grossepunctatus Reitter 1905
Onthophagus guatemalensis Bates 1888
Onthophagus hecate (Panzer) 1794
Onthophagus hecate blatchleyi Brown 1929
Onthophagus herus Peringuey 1901
Onthophagus hildebrandti Harold 1878
Onthophagus hirticulus d'Orbigny 1915
Onthophagus hopfneri Harold 1869
Onthophagus hystrix Boucomont 1914
Onthophagus illyricus (Scopoli) 1763
Onthophagus incanus MacLeay 1888
Onthophagus incensus Say 1835
Onthophagus insignis Peringuey 1896
Onthophagus interstitialis Fahraeus 1857
Onthophagus jangga Matthews 1972
Onthophagus jeaneli d'Orbigny 1913
Onthophagus kchatriya Boucomont 1914
Onthophagus kiambram Storey 1977
Onthophagus knulli Howden & Cartwright 1963
Onthophagus lacustris Harold 1877
Onthophagus laevigatus (Fabricius) 1798
Onthophagus lamelliger Gerstaecker 1871
Onthophagus laminatus MacLeay 1864
Onthophagus landolti Harold 1880
Onthophagus latestriatus d'Orbigny 1908
Onthophagus lebasi Boucomont 1932
Onthophagus lemur (Fabricius) 1781
Onthophagus lenzi Harold 1874
Onthophagus macrocephalus Kirby 1818
Onthophagus maki (Illiger) 1803
Onthophagus marginicollis Harold 1880
Onthophagus medorensis Brown 1929
Onthophagus mexicanus Bates 1887
Onthophagus mirabilis Bates 1887
Onthophagus mniszechi Harold 1869
Onthophagus mucronatus Thomson 1858
Onthophagus mulgravei Paulian 1937
Onthophagus mundill Matthews 1972
Onthophagus neostenocerus Goidanich 1926
Onthophagus nigricornis Fairmaire 1887
Onthophagus nitidior Bates 1887
Onthophagus nitidulus Klug 1845
Onthophagus notatus d'Orbigny 1902
Onthophagus nuchicornis (Linnaeus) 1758
Onthophagus nurubum Matthews 1972
Onthophagus nyctopus Bates 1887
Onthophagus obliquus (Olivier) 1789
Onthophagus obtusicornis Fahraeus 1857
Onthophagus ocellifer d'Orbigny 1902
Onthophagus ohbayashii Nomura
Onthophagus oklahomensis Brown 1927
Onthophagus orientalis Harold 1868
Onthophagus orpheus canadensis (Fabricius) 1801
Onthophagus ovatus (Linnaeus) 1767
Onthophagus pallidipennis Fahraeus 1857
Onthophagus paluma Matthews 1972
Onthophagus pardalis (Fabricius) 1798
Onthophagus parryi Harold
Onthophagus parumnotatus Fahraeus 1857
Onthophagus pennsylvanicus Harold 1873
Onthophagus plebejus Klug 1855
Onthophagus praecellens Bates 1887
Onthophagus producticollis d'Orbigny 1908
Onthophagus pseudocervulus Reitter
Onthophagus pugionatus Fahraeus 1857
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pygargus Motschulsky 1845
Onthophagus pygmaeus (Schaller) 1783
Onthophagus quadraticeps Harold 1867
Onthophagus quadricuspis d'Orbigny 1908
Onthophagus quadridentatus (Fabricius) 1798
Onthophagus quadrinodosus Fahraeus 1857
Onthophagus raffreyi Harold 1886
Onthophagus ramosus Wiedemann 1823
Onthophagus rasipennis d'Orbigny 1908
Onthophagus recticornutus Lansberge 1883
Onthophagus retusus Harold 1869
Onthophagus rhinolophus Harold 1869
Onthophagus rhinophyllus Harold 1868
Onthophagus rubefactus d'Orbigny 1913
Onthophagus rubrimaculatus MacLeay 1864
Onthophagus sellatus Klug 1893
Onthophagus semiflavus Boheman 1860
Onthophagus semimetallicus Lea 1923
Onthophagus sharpi Harold 1875
Onthophagus sidneyensis Blackburn 1903
Onthophagus signatus Fahraeus 1857
Onthophagus similis (Scriba) 1790
Onthophagus simpliciceps d'Orbigny 1905
Onthophagus sloanei Blackburn 1903
Onthophagus socialis Arrow 1931
Onthophagus spinifex (Fabricius) 1781
Onthophagus stockwelli Howden & Young 1981
Onthophagus striatulus (Beauvois) 1805
Onthophagus subaeneus Beauvois 1805
Onthophagus subopacus Robinson 1940
Onthophagus sugillatus Klug 1855
Onthophagus tarandus (Fabricius) 1792
Onthophagus taurus (Schreber) 1759
Onthophagus tenebrosus Harold 1871
Onthophagus transcaspicus Koenig 1888
Onthophagus truchmenus Kolenati 1846
Onthophagus truncaticornis Schaller 1783
Onthophagus tuberculatus (Waterhouse) 1874
Onthophagus tuberculifrons Harold 1873
Onthophagus turbatus Walker 1858
Onthophagus tweedensis Blackburn 1901
Onthophagus unifasciatus Schaller 1783
Onthophagus vacca (Linnaeus) 1767
Onthophagus variegatus (Fabricius) 1798
Onthophagus variegranosus d'Orbigny 1905
Onthophagus velutinus Horn 1875
Onthophagus vinctus Erichson 1843
Onthophagus virescens Erichson 1843
Onthophagus vitulus Fabricius 1776
Onthophagus vividus Arrow 1907
Onthophagus vylderi d'Orbigny 1913

Oruscatus davus (Erichson) 1847

Oxysternon conspicillatum (Weber) 1801
Oxysternon festivum (Linnaeus)
Oxysternon macleayi (Nevinson) 1892

Pachylomera femoralis Kirby 1828

Parapinotus beccarii (Harold) 1875
Parapinotus dewitzi Harold 1878

Pedaria armata Raffray 1877

Pedaridium pilosum (Robinson) 1948

Phalops ardea Klug 1855
Phalops aurifrons Fairmaire 1884
Phalops boschas Klug 1855
Phalops divisus Lansberge 1883
Phalops dregei Harold 1867
Phalops flavocinctus Klug 1855
Phalops prasinus Erichson 1843
Phalops smaragdinus Harold 1875
Phalops vanellus Lansberge 1883
Phalops wittei Harold 1867

Phanaeus amethystinus amethystinus Harold 1863
Phanaeus amithaon Harold
Phanaeus chalcomelas (Perty) 1830
Phanaeus daphnis Harold 1863
Phanaeus demon Laporte 1840
Phanaeus difformis LeConte 1847
Phanaeus endymion Harold 1863
Phanaeus furiosus Bates 1887
Phanaeus guatemalensis Harold 1871
Phanaeus haroldi Kirsch 1871
Phanaeus howdeni Arnaud 1983
Phanaeus igneus MacLeay 1819
Phanaeus kirbyi Vigors 1825
Phanaeus meleagris Blanchard 1843
Phanaeus mexicanus Harold 1863
Phanaeus nimrod Harold 1863
Phanaeus obliquens Bates 1889
Phanaeus palaeno Blanchard 1843
Phanaeus palliatus Sturm 1883
Phanaeus prasinus Harold 1868
Phanaeus pyrois Bates 1887
Phanaeus quadridens (Say) 1835
Phanaeus splendidulus (Fabricius) 1781
Phanaeus triangularis Say 1823
Phanaeus tridens pseudofurcosus Balthasar
Phanaeus vindex MacLeay 1819
Phanaeus wagneri pilatei Harold 1863
Phanaeus wagneri wagneri Harold 1863

Proagoderus aciculatus (Fahraeus) 1857
Proagoderus alcyonides (d'Orbigny) 1913
Proagoderus aureiceps (d'Orbigny) 1902
Proagoderus bicallosus (Klug) 1855
Proagoderus brucei (Reiche) 1847
Proagoderus chrysopes (Bates) 1888
Proagoderus mouhoti (Harold) 1875
Proagoderus rangifer (Klug) 1855
Proagoderus speculicollis (Quedenfeldt) 1884
Proagoderus tersidorsis (d'Orbigny) 1902

Pseudocanthon perplexus (LeConte) 1847

Sarophorus costatus (Fahraeus) 1857

Scarabaeus aegyptiorum Latreille 1827
Scarabaeus ambiguus (Boheman) 1857
Scarabaeus andersoni (Felsche) 1907
Scarabaeus benningseni (Felsche) 1907
Scarabaeus bohemani Harold 1868
Scarabaeus canaliculatus Fairmaire 1888
Scarabaeus catenatus Gerstaecker 1871
Scarabaeus cicatrosus (Lucas) 1846
Scarabaeus cristatus Fabricius 1775
Scarabaeus devotus Redtenbacher 1848
Scarabaeus erichsoni Harold 1867
Scarabaeus flavicornis (Boheman) 1860
Scarabaeus galenus (Westwood) 1847
Scarabaeus gariepinus (Ferreira) 1953
Scarabaeus goryi Laporte 1840
Scarabaeus hippocrates MacLeay 1821
Scarabaeus laevistriatus Fairmaire 1893
Scarabaeus laticollis Linnaeus 1767
Scarabaeus parvulus (Boheman) 1860
Scarabaeus proboscideus (Guerin) 1844
Scarabaeus radama Fairmaire 1895
Scarabaeus rotundigena (Felsche) 1907
Scarabaeus rubripennis (Boheman) 1860
Scarabaeus rugosus (Hausmann) 1807
Scarabaeus rusticus (Boheman) 1857
Scarabaeus sacer Linnaeus 1758
Scarabaeus satyrus (Boheman) 1860
Scarabaeus semipunctatus (Fabricius) 1792
Scarabaeus striatus (Laporte) 1840
Scarabaeus variolosus (Fabricius) 1787
Scarabaeus westwoodi Harold 1869
Scarabaeus zambesianus

Scatimus ovatus Harold 1862
Scatimus quadridentatus Balthasar 1939

Sceliages adamastor Serville 1828
Sceliages brittoni Zur Strassen 1963

Sisyphus goryi Harold 1859
Sisyphus hirtus Wiedemann 1823
Sisyphus impressipennis Lansberge 1886
Sisyphus longipes (Olivier) 1789
Sisyphus mexicanus Harold 1863
Sisyphus muricatus Olivier 1789
Sisyphus schaefferi Linnaeus 1758

Sphaerocanthon clypeatus (Laporte) 1840

Stereopyge douei (Gory) 1891

Sulcophanaeus batesi (Harold) 1868
Sulcophanaeus carnifex (Linnaeus) 1758
Sulcophanaeus cupricollis (Nevinson) 1891
Sulcophanaeus imperator (Chevrolat) 1844
Sulcophanaeus leander (Waterhouse) 1891
Sulcophanaeus menelas (Castelnau) 1840
Sulcophanaeus noctis (Bates) 1887
Sulcophanaeus velutinus (Murray) 1856

Temnoplectron politulum MacLeay 1887
Temnoplectron rotundum Westwood 1841
Temnoplectron subvolitans Matthews 1974

Tiniocellis spinipes (Roth) 1851

Tragiscus dimidiatus Klug 1855

Troxys bahianus Boucard 1828

Uroxys bahianus Boucomont 1928
Uroxys bidentis Howden & Young 1981
Uroxys depressifrons Howden & Young 1981
Uroxys dybasi Howden & Young 1981
Uroxys gorgon Arrow 1933
Uroxys macrocularis Howden & Young 1981
Uroxys metagorgon Howden & Young 1981
Uroxys micros Bates 1887
Uroxys platypyga Howden & Young 1981
Uroxys transversifrons Howden & Gill 1981

Xinidium dentilabris Harold 1869