Gordon Hendler: Publications and downloads
Hendler, G., A.E. Migotto, C.R.R. Ventura, and L. Wilk (2012) Epizoic Ophiothela brittle stars have
invaded the Atlantic. Coral Reefs. DOI 10.1007/s00338-012-0936-6. PDF
invaded the Atlantic. Coral Reefs. DOI 10.1007/s00338-012-0936-6. PDF
Hendler, G. (2011) New light on the nomenclature, taxonomy, and biology of Hemipholis species
(Echinodermata: Ophiuroidea: Ophiactidae). Zootaxa 3048: 44-52.
(Echinodermata: Ophiuroidea: Ophiactidae). Zootaxa 3048: 44-52.
Hendler, G. and I.-H Kim (2010) Larval biology of thaumatopsyllid copepods endoparasitic in
Caribbean ophiuroids. Journal of Crustacean Biology 30: 206-224.
Hendler, G. and M. Dojiri (2009) The contrariwise life of a parasitic, pedomorphic copepod with a
non-feeding adult: ontogenesis, ecology, and evolution. Invertebrate Biology 128: 65-82.
Dojiri, M., G. Hendler, and I.-H Kim (2008) Larval development of Caribeopsyllus amphiodiae
(Thaumatopsyllidae, Copepoda), an enterozoic parasite of the brittle star Amphiodia urtica.
Journal of Crustacean Biology 28: 281-305.
Hendler, G. (2007) Class Ophiuroidea, Pp. 930-941. In: J.T. Carlton (ed.) The Light and Smith
Manual: Intertidal Invertebrates of Central California and Oregon (Fourth Edition).
University of California Press, Berkeley.
Hendler, G. (2005) Two new brittle star species of the genus Ophiothrix (Echinodermata:
Ophiuroidea: Ophiotrichidae) from coral reefs in the southern Caribbean Sea, with notes
on their biology. Caribbean Journal of Science 41: 583-599.
Hendler, G. (2004) An echinoderm’s eye view of photoreception and vision. Pp. 339-349. In: T.
Heinzeler and J.H. Nebelsick (eds.) Echinoderms: München. A.A. Balkema, Leiden.
Mallefet, J., G. Hendler, C.M. Herren, C. McDougall and J. Case (2004) A new bioluminescent
ophiuroid species from the coast of California. Pp. 305-310. In: T. Heinzeler and J.H.
Nebelsick (eds.) Echinoderms: München. A.A. Balkema, Leiden.
Aizenberg, J. and G. Hendler (2004) Designing efficient microlens arrays: lessons from nature.
Journal of Materials Chemistry 14: 2066-2072.
Hendler, G. (2003) Canary of the sea, Beauty in the deep, Hancock’s Velero discoveries, Seeing
stars. Pp. 16, 69, 70 and 74. In: B. Binet and E. Bailey (eds.) 90 Years, 90 Treasures.
Natural History Museum of Los Angeles County. Los Angeles, California.
Ho, J.-S., M. Dojiri, G. Hendler, and G.B. Deets (2003) A new species of Copepoda
(Thaumatopsyllidae) associated with a brittle star from California, USA, and designation of
a new Order Thaumatopsylloida. Journal of Crustacean Biology 23: 582-594.
Hendler, G. (2002) Ophiuroidea. Pp. 414-416. In: J.W. Nybakken, W.W. Broenkow, T.L. Vallier
(eds.) Interdisciplinary Encyclopedia of Marine Sciences. Grolier Academic Reference,
Danbury, Connecticut.
Hendler, G. (2002). Marine Fauna from Isla San Felix, Chile. Pp. 249-256. In: R.W. Schmieder (ed.)
XROX: The 2002 San Felix Island Expedition. Cordell Expeditions, 4295 Walnut Blvd.,
Walnut Creek, California 94596.
Hendler, G. (2002) Echinoderms. Pp. 282-287. In: N. Eldredge (ed.) Life on earth. An
encyclopedia of biodiversity, ecology, and evolution. Volume 1 A-G. ABC Clio, Santa
Barbara, California.
Hendler, G. (2002) Account of Ophionereis diabloensis, a new species of brittle star, and O.
amphilogus, with information on their brooding reproduction and distribution
(Echinodermata: Ophiuroidea: Ophionereididae). Proceedings of the Biological Society
of Washington 115: 57-74.
Hendler, G. (2001) That’s no worm… (Letter) Science 291: 47.
Hendler, G. and C.J. Bundrick (2001) A new, brooding brittle star from California (Echinodermata:
Ophiuroidea). Contributions in Science, Natural History Museum of Los Angeles County
No. 486: 1-11.
Hendler, G. and L.U. Tran. (2001) Reproductive biology of a deep-sea brittle star Amphiura
carchara (Echinodermata: Ophiuroidea). Marine Biology 138: 113-123.
Aizenberg, J., A.Tkachenko, S.Weiner, L. Addadi, and G. Hendler (2001) Calcitic microlenses as
part of the photoreceptor system in brittlestars. Nature 412: 819-822.
Fujita, T. and G. Hendler (2001) Description of a new species of Astrophiura (Echinodermata:
Ophiuroidea) from Tosa Bay, Japan, and several of its remarkable anatomical
characteristics. Pp. 263-281. In: T. Fujita, H, Saito and M. Takeda (eds.) Deep Sea Fauna
and Pollutants in Tosa Bay, National Science Museum Monographs, No. 20, Tokyo.
Hendler, G. (2000) Forward. Pp. 7-9. In: Schoppe, S. A guide to common shallow water sea
stars, brittle stars, sea urchins, sea cucumbers and feather stars (Echinoderms) of the
Philippines. Times Media Private Limited, Singapore.
Hendler, G. (2000) Babysitting brittle-stars, kidnapping cuckoos, and other deceptions in the
nursery. Terra 37: 8-15.
Hendler, G. and D.L. Pawson (2000) The echinoderms of the rhomboidal cays, Belize:
biodiversity, distribution, and ecology. Atoll Research Bulletin No. 479: 273-299.
Humes, A.G. and G. Hendler (1999) Biology and taxonomy of species of Ophiopsyllus and
Pseudanthessius (Copepoda) associated with brittle stars (Ophiuroidea) in Belize.
Bulletin of Marine Science 65: 699-713.
Hendler, G., C.C. Baldwin, D.G. Smith, and C.E. Thacker (1999) Planktonic dispersal of juvenile
brittle stars (Echinodermata: Ophiuroidea) on a Caribbean reef. Bulletin of Marine
Science 65: 283-288.
Hendler, G., M.J. Grygier, E. Maldonado, and J. Denton (1999) Babysitting brittle stars:
Heterospecific symbiosis between ophiuroids (Echinodermata). Invertebrate Biology
118: 190-201.
Hendler, G. (1998) Implications of the remarkable ontogenetic changes in some deep-sea brittle
stars. Pp. 353-358. In: R. Mooi and M. Telford (eds.) Echinoderms: San Francisco.
Balkema, Rotterdam.
Pearse, V.B., J.S. Pearse, G. Hendler, and M. Byrne (1998) Discovery of an accessible
population of Ophiocanops, off NE Sulawesi, Indonesia. Pp. 413-418. In: R. Mooi and M.
Telford (eds.) Echinoderms: San Francisco. Balkema, Rotterdam.
Hendler, G. (1996) Crinoidea. Pp. 85-95. In: J.A. Blake and A. Lissner (eds.) Taxonomic Atlas of
the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Santa
Barbara Museum of Natural History.
Hendler, G. (1996) Ophiuroidea. Pp. 113-176. In: J.A. Blake and A. Lissner (eds.) Taxonomic
Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara
Channel. Santa Barbara Museum of Natural History.
Hendler, G. (1996) Echinodermata collected at Rocas Alijos. Pp. 319-338. In: R.W.
Schmieder (ed.) Rocas Alijos. Kluwer Academic Publishers, Dordrecht, Netherlands.
Dearborn, J. H., G. Hendler, and K. C. Edwards (1996) The diet of Ophiosparte gigas
(Echinodermata: Ophiuroidea) along the Antarctic Peninsula, with comments on its
taxonomic status. Polar Biology 16: 309-320.
Hendler, G., J.E. Miller, D.L. Pawson, and P.M. Kier (1995) Sea stars, sea urchins, and their
allies: Echinoderms of Florida and the Caribbean. Smithsonian Institution Press. 390 pp.
Hendler, G. (1995) Underwater expedition to study brittle stars. Terra 32:6.
Hendler, G. (1995) New species of brittle stars from the western Atlantic: Ophionereis vittata,
Amphioplus sepultus, and Ophiostigma siva, and the designation of a neotype for
Ophiostigma isocanthum (Say) (Echinodermata: Ophiuroidea). Contributions in
Science, Natural History Museum of Los Angeles County 458: 1-19.
Hendler, G. (1991) Stars of the Florida Keys. Terra 30: 26‑33.
Hendler, G. (1991) Chapter 6 ‑ Echinodermata: Ophiuroidea. Pp. 355‑511. In: A.C. Giese and J.S.
Pearse (eds.) Reproduction of Marine Invertebrates, Vol. VI. Blackwell Scientific and The
Boxwood Press, Pacific Grove, California.
Hendler, G. and J. E. Miller (1991) Swimming ophiuroids ‑ real and imagined. Pp. 179‑190. In:
Yanagisawa, T., I. Yasumasu, C. Oguro, N. Suzuki, and T. Motokawa (eds.) Biology of
Echinodermata. A.A. Balkema, Rotterdam, Netherlands.
Hendler, G. and J.H. Dearborn (1990) Ophiuroidea (Schlangensterne). Pp. 176-181. In: W.
Wagele and J. Sieg (eds.) Fauna der Antarktis. J.Paul Parey Publishers. [in German].
Cobb, J.L.S. and G. Hendler (1990) Neurophysiological characterization of the photoreceptor
system in a brittlestar, Ophiocoma wendtii (Echinodermata: Ophiuroidea). Comparative
Biochemistry and Physiology. 97A: 329‑333.
Byrne, M. and G. Hendler (1988) Arm structures of the ophiomyxid brittlestars (Echinodermata:
Ophiuroidea: Ophiomyxidae). Pp. 687‑695. In: Burke, R.D., P.V. Mladenov, P. Lambert,
and R.L. Parsley (eds.) Echinoderm Biology. A.A. Balkema, Rotterdam, Netherlands.
Hendler, G. (1988) Ophiuroid skeleton ontogeny reveals homologies among skeletal plates of
adults: A study of Amphiura filiformis, Amphiura stimpsonii and Ophiophragmus
filograneus (Echinodermata). Biological Bulletin 174: 20‑29.
Hendler, G. (1988) Western Atlantic Ophiolepis (Echinodermata: Ophiuroidea): a description of
O. pawsoni, n. sp., and a key to the species. Bulletin of Marine Science 42: 265‑272.
Hendler, G. and D. L. Pawson (1988) Echinoderm nomenclature: Not in need of repair.
Systematic Zoology 36: 395‑396.
Hendler, G. and R.W. Peck (1988) Ophiuroids off the deep end: Fauna of the Belizean fore‑reef
slope. Pp. 411‑419. In: Burke, R.D., P.V. Mladenov, P. Lambert, and R.L. Parsley (eds.)
Echinoderm Biology. A.A. Balkema, Rotterdam, Netherlands.
Hendler, G. and M. Byrne (1987) Fine structure of the dorsal arm plate of Ophiocoma wendti:
Evidence for a photoreceptor system (Echinodermata: Ophiuroidea). Zoomorphology
107: 261‑272.
Hendler, G. and R.L. Turner (1987) Two new species of Ophiolepis (Echinodermata:
Ophiuroidea) from the Caribbean Sea and Gulf of Mexico: with notes on ecology,
reproduction and morphology. Contributions in Science, Natural History Museum of
Los Angeles County 395: 1‑14.
Hendler, G. and B.S. Littman (1986) The ploys of sex: relationships among the mode of
reproduction, body size and habitats of coral reef brittlestars. Coral Reefs 5: 31‑42.
Hendler, G. and P. Tyler (1985) The reproductive cycle of Ophioderma brevispinum
(Echinodermata: Ophiuroidea). P.S.Z.N.I. Marine Ecology 7:115‑122.
Hendler, G. (1984) Brittlestar color‑change and phototaxis (Echinodermata: Ophiuroidea:
Ophiocomidae). P.S.Z.N.I. Marine Ecology 5:379‑401.
Hendler, G. and J.E. Miller (1984) Ophioderma devaneyi and Ophioderma ensiferum, new
brittlestar species from the Western Atlantic (Echinodermata: Ophiuroidea). Proceedings
of the Biological Society of Washington 97: 442‑461.
Hendler, G. and J. E. Miller (1984) Feeding behavior of Asteroporpa annulata, a gorgonocephalid
brittlestar with unbranched arms. Bulletin of Marine Science 34: 449‑460.
Hendler, G. (1984) The association of Ophiothrix lineata and Callyspongia vaginalis: A
brittlestar‑sponge cleaning symbiosis? P.S.Z.N.I. Marine Ecology 5: 9‑27.
Hendler, G. (1982) An echinoderm vitellaria with a bilateral larval skeleton: Evidence for the
evolution of ophiuroid vitellariae from ophioplutei. Biological Bulletin 163: 431‑437.
Hendler, G. (1982) Slow flicks show star tricks: Elapsed‑time analysis of basketstar
(Astrophyton muricatum) feeding behavior. Bulletin of Marine Science 32: 909‑918.
Hendler, G. (1982) The feeding biology of Ophioderma brevispinum (Echinodermata:
Ophiuroidea). Pp. 21‑27. In: J.M. Lawrence (ed.) Proceedings of the International
Echinoderms Conference ‑ Tampa Bay. A.A. Balkema, Rotterdam.
Hendler, G. and D.R. Franz (1982) The biology of a brooding seastar, Leptasterias tenera, in
Block Island Sound. Biological Bulletin 162: 273‑289.
Hendler, G. and D.L. Meyer (1982) Ophiuroids flagrante delicto and notes on the spawning
behavior of other echinoderms in their natural habitat. Bulletin of Marine Science 32:
600‑607.
Hendler, G. and D. L. Meyer (1982) An association of a polychaete, Branchiosyllis exilis with an
ophiuroid, Ophiocoma echinata, in Panama. Bulletin of Marine Science 32: 736‑744.
Dearborn, J.H., L.E. Watling, K.C. Edwards, D.B. Fratt and G. Hendler (1982) Echinoderm biology
and general benthic collecting along the Antarctic Peninsula. Antarctic Journal of the
United States 17: 162‑164.
Hendler, G. (1979) Sex‑reversal and viviparity in Ophiolepis kieri n.sp. with notes on viviparous
brittlestars from the Caribbean (Echinodermata: Ophiuroidea). Proceedings of the
Biological Society of Washington 9: 783‑795.
Hendler, G. (1979) Reproductive periodicity of ophiuroids (Echinodermata: Ophiuroidea) on the
Atlantic and Pacific Coasts of Panama. Pp. 145‑156. In: S. Stancyk (ed.) Reproductive
Ecology of Marine Invertebrates. Belle W. Baruch Library in Marine Science No. 9.
University of South Carolina, Columbia.
Hendler, G. (1978) Development of Amphioplus abditus (Verrill) (Echinodermata: Ophiuroidea). II.
Description and discussion of ophiuroid skeletal ontogeny and homologies. Biological
Bulletin 154: 79‑95.
Hendler, G. (1977) The differential effects of seasonal stress and predation on the stability of
reef‑flat echinoid populations. Proceedings of the Third International Coral Reef
Symposium I: 217‑223.
Worley, E.K., D.R. Franz and G. Hendler (1977) Seasonal patterns of gametogenesis in a North
Atlantic brooding asteroid, Leptasterias tenera. Biological Bulletin 153: 237‑253.
Hendler, G. (1977) Development of Amphioplus abditus (Verrill) (Echinodermata: Ophiuroidea). I.
Larval biology. Biological Bulletin 152: 51‑63.
Hendler, G. (1975) Adaptational significance of the patterns of ophiuroid development. American
Zoologist 15: 691‑715.
Hendler, G. (1973) Northwest Atlantic amphiurid brittlestars, Amphioplus abditus (Verrill),
Amphioplus macilentus (Verrill), and Amphioplus sepultus n. sp. (Ophiuroidea:
Echinodermata): Contributions on their systematics, zoogeography, annual periodicity, and
larval adaptations. Ph. D. Thesis. University of Connecticut. 254 pp.
Humes, A.G. and G. Hendler (1972) New cyclopoid copepods associated with the ophiuroid
genus Amphioplus on the eastern coast of the United States. Transactions of the
American Microscopical Society 91: 539‑555.
Hendler, G. and D. R. Franz (1971) Population dynamics and life history of Crepidula convexa
Say (Gastropoda: Prosobranchia) in Delaware Bay. Biological Bulletin 141: 514‑526.
Franz, D.R. and G. Hendler (1970) Substrate diversity and taxonomy of Crepidula convexa (Say)
(Gastropoda: Prosobranchia). Occasional Papers of the University of Connecticut
(Biology) 1: 289‑291.
Loveland, R.E., G. Hendler, and G. Newkirk (1969) New records of nudibranchs from New
Jersey. Veliger 11: 418‑420.